Upsilon is the twentieth Greek letter, from Phoenician waw; its value is 400. Upsilon has a genuinely attested moral symbolism as the 'Pythagorean letter' (littera Pythagorica): its forking shape was read as an emblem of the crossroads of life, the point at which the young must choose between the paths of virtue and of vice.
The Greek letters doubled as numerals in the Milesian (alphabetic) system, in which each letter carried a fixed value; the practice of summing a word's letters into a number, called isopsephy, is the Greek counterpart of Hebrew gematria and appears in inscriptions, the Greek Magical Papyri and early Christian texts. Upsilon's Phoenician ancestry places it in a borrowing whose exact point of contact, somewhere among Al Mina, Cyprus, Crete and Euboea in the Aegean world, is still disputed, though the borrowing itself, conventionally dated to around the 8th century BCE, is agreed to be the ancestor of every European alphabet still in use, including the Latin one covered elsewhere in this atlas.
The Greeks' own contribution was to give vowels dedicated letters of their own, something the consonant-only Phoenician script never did. Its isopsephy value of 400 belongs to the Milesian alphabetic numeral system, which also needed three further, otherwise obsolete letters, digamma (also called stigma), koppa and sampi, to cover every value from 1 up to 900; none of the three survives among the 24 letters in ordinary use today.